Darling Marine Center
193 Clarks Cove Road
Walpole, ME 04573

207-563-3146
207-563-3119 (fax)

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Dr. Damian Brady Dr. Pete Jumars Dr. Bob Steneck
Annette deCharon Dr. Larry Mayer Dr. Rick Wahle
Dr. Kevin Eckelbarger Dr. Mary Jane Perry Dr. Rhian Waller
Dr. Lewis Incze Dr. Warren Riess Dr. Les Watling

Dr. Damian C. Brady
Assistant Research Professor
Ph.D. University of Delaware, 2008

damian.brady@maine.edu
more info

Research Interests: Water quality modeling, individual-based modeling, behavioral ecology, and ecophysiology of fish

The spatial and temporal dynamics of water quality and organism behavior determine exposure and dictate the extent to which poor water quality impacts habitat function. My work focuses on making the connection between water quality and living resources using numerical models that incorporate water quality monitoring and the ecophysiology of the affected organisms.

  • McMahan, M.D., Brady, D.C., Cowan, D.F., Grabowski, J.H., & Sherwood, G.D. (accepted with major revisions) Using fine-scale acoustic telemetry to observe the effects of a groundfish predator (Atlantic cod, Gadus morhua) on the movement behavior of the American lobster (Homarus americanus). Canadian Journal of Fisheries and Aquatic Sciences.
  • Testa, J.M., Brady, D.C., Di Toro, D.M., Boynton, W.R., Cornwell, J.C., & Kemp, W.M. (2013) Sediment Flux Modeling: Simulating nitrogen, phosphorus, and silica cycles. Estuarine, Coastal and Shelf Science doi:10.1016/j.ecss.2013.06.014.
  • Brady, D.C. & Targett, T.E. (2013) Movement of juvenile weakfish (Cynoscion regalis) and spot (Leiostomus xanthurus) in relation to diel-cycling hypoxia in an estuarine tributary: Assessment using acoustic telemetry. Marine Ecology Progress Series doi:10.3354/meps10466.
  • Zhang, Q., Brady, D.C., & Ball, W.P. (2013) Long-term seasonal trends of nitrogen, phosphorus, and suspended sediment load from the non-tidal Susquehanna River Basin to Chesapeake Bay. Science of the Total Environment, 452-453: 208-221.
  • Brady, D.C., Testa, JM., Di Toro, D.M., Boynton, W.R., Kemp, W.M. (2013) Sediment Flux Modeling: Application and validation for coastal systems. Estuarine, Coastal, and Shelf Science 117: 107-124.
  • Grieve, C., Brady, D.C., Polet, H. 2011 Best Practices for Managing, Measuring, and Mitigating the Benthic Impacts of Fishing: Final Report to the Marine Stewardship Council.
  • Brady, D. C. and T. E. Targett (2010). Characterizing the escape response of juvenile summer flounder Paralichthys dentatus to diel-cycling hypoxia. Journal of Fish Biology 77(1): 137-152.
  • Fennel, K., Brady, D.C., Di Toro, D.M., Fulweiler, R., Gardner, W.S., Giblin, A., McCarthy, M.J., Rao, A., Seitzinger, S., Thouvenot-Korppoo, Tobias, C. (2009) Modeling denitrification in aquatic sediments. Biogeochemistry. 93(1-2): 159-178.
  • Brady, D.C., Tuzzolino, D.M., Targett, T.E. (2009) Behavioral responses of juvenile weakfish, Cynoscion regalis, to diel-cycling hypoxia: swimming speed, angular correlation, expected displacement and effects of hypoxia acclimation. Canadian Journal of Fisheries and Aquatic Sciences. 66(3): 415-424.
  • Tyler, R.M., Brady, D.C., Targett, T.E. (2009) Temporal and spatial dynamics of diel-cycling dissolved oxygen in estuarine tributaries. Estuaries and Coasts. 32(1): 123-145.
  • Breitburg, D.L., Craig, J.K., Fulford, R.S., Rose, K.A., Boynton, W.R., Brady, D.C., Ciotti, B.J., Diaz, R.J., Friedland, K.D., Hagy, J.D. III, Hart, D.R., Hines, A.H., Houde, E.D., Kolesar, S.E., Nixon, S.W., Rice, J.A., Secor, D.H., Targett, T.E. (2009) Nutrient enrichment and fisheries exploitation: interactive effects on estuarine living resources and their management. Hydrobiologia, 629(1): 31-47.
  • CBEO Project Team: Ball, W.P., Brady, D.C., Brooks, M.T., Burns, R, Cuker, B.E., Di Toro, D.M., Gross, T.F., Kemp, W.M., Murray, L., Murphy, R.R., Perlman, E., Piasecki, M., Testa, J.M., Zaslavsky, I. (2008) Prototype system for multi-disciplinary shared cyberinfrastructure: Chesapeake Bay Environmental Observatory (CBEO). Journal of Hydrologic Engineering, ASCE. 13(10): 960-970.

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Annette deCharon

Annette deCharon
Senior Marine Outreach Education Scientist
Director, CODEE-OS
M.S. Oceanography, Oregon State University, 1988

annette.decharon@maine.edu
more info
COSEE-OS

Research Interests: Education, Oceanography, Marine Policy

I am particularly interested in developing innovative multimedia for teaching and learning about ocean systems. Since 1997, I have authored over 30 web-based and other multimedia publications including the award-winning "Phytopia: Discovery of the Marine Ecosystem" CD-ROM. I am currently Director of the Centers for Ocean Sciences Education Excellence - Ocean Systems (COSEE-OS). Our long-term goal is to help the COSEE Network reach rural and inland audiences. Although located on the coast of Maine, I am happy to continue my association with NASA as education lead for the upcoming Aquarius mission (2011 launch) which will measure global ocean surface salinity.

  • A. deCharon, 2010. COSEE-Ocean Systems: Collaborative Mapping to Clarify Concepts in Ocean Observing. The Earth Scientist: Journal of the National Earth Science Teachers Association, Vol. 26(1), 29-30.
  • A. deCharon, 2010. Seeking Salt: NASA Aquarius to Fill Void in Ocean-Climate Understanding. The Earth Scientist: Journal of the National Earth Science Teachers Association, Vol. 26(1), 14-16.
  • A. deCharon, J. Albright, C. Herren, A. Cline, and J. Repa. 2009. Online Tools Help Get Scientists and Educators on the Same Page. Eos, Transactions, American Geophysical Union, 90(34): 289-290
  • G. Lagerloef, F. Colomb, D. Le Vine, F. Wentz, S. Yueh, C. Ruf, J. Lilly, J. Gunn, Y. Chao, A. deCharon, G. Feldman, and C. Swift. 2008. The Aquarius / SAC-D Mission: Designed to Meet the Salinity Remote-Sensing Challenge," Oceanography, v. 21(1), 68-81.
  • A. deCharon, S. Etheridge, T. McDonnell Wysor, and M. Sieracki, 2006. HAB outreach using multimedia: integrating ocean research and education. African Journal Marine Science, 28(2): 459-463
  • A. deCharon and J. Ryan. 2006. Phytopia: Upwelling Zones. URL: www.bigelow.org/upwelling
  • A. deCharon and M. Sieracki. 2003. Phytopia: Discovery of the Marine Ecosystem. educational CD-ROM
  • A. deCharon, M. Sieracki and P. Wilson. 2002. Cycling through the Food Web. URL: www.bigelow.org/bacteria
  • T. Wysor, A. deCharon, and R. Wahle. 2004. Hatch to Catch II... the Journey Continues. URL: www.bigelow.org/h2c2/
  • A. deCharon and S. Etheridge. 2001. Toxic & Harmful Algal Blooms. URL: www.bigelow.org/hab/
  • A. deCharon, L. Incze, R. Wahle and N. Wolff. 2000. Hatch to Catch. URL: www.bigelow.org/hatch_to_catch
  • A. deCharon and M. Sieracki. 1999. Ship Mates: Explore the Gulf of Maine as Oceanographers Do. URL: www.bigelow.org/shipmates

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Kevin Eckelbarger

Dr. Kevin Eckelbarger
Professor, School of Marine Sciences &
Ph.D. Northeastern University, 1974

kevine@maine.edu
more info

Research interests: reproductive and developmental biology of marine invertebrates; deep-sea biology

I have been conducting long-term research on the factors controlling reproductive cycles in marine invertebrates, particularly those in bathyal and deep-water habitats. Much of my work involves the use of oceanographic research ships and manned submersibles. For the past 20 years, most of this research has been conducted in the Gulf of Mexico, the Bahama Islands, and the Gulf of Maine in collaboration with colleagues at the University of Southampton (Britain) and the Oregon Institute of Marine Biology. A good deal of my work focuses on the ultrastructural features of gametogenesis, especially oogenesis, and the mechanisms of yolk synthesis in the egg. Through the use of electron microscopy, I have been able to better understand how food drives egg production in deep water invertebrates that often have access to limited food supplies. Of particular interest are the factors that determine whether a deep sea species undergoes annual or continuous reproduction, often in the absence of environmental cues (light, food, temperature change) that influence the reproductive cycles of shallow water species.

  • Hodgson, A.H., K.J. Eckelbarger, C.M. Young. 2009. Sperm ultrastructure and spermatogenesis in the hydrothermal vent gastropod Rhychopelta concentrica. Journal Molluscan Studies 75: 159-165.
  • Blake, J.A., J.P. Grassle, K.J. Eckelbarger, 2009. Capitella teleta, a new species designation for the opportunistic and experimental Capitella sp.I with a review of the literature and confirmed records. Zoosymposia 2: 25-54.
  • Mosher, C.V., L. Watling, K.J. Eckelbarger. 2009. Deep-ea Ophiuroidea associated with seamount communities of the Western North Atlantic. Proceedings of the 12th International Echinoderm Conference, pp. 235-239.
  • Eckelbarger, K.J., C. Hand, and K. Uhlinger. 2008. Ultrastructural features of trophonema formation and oogenesis in the starlet sea anemone, Nematostella vectensis (Edwardsiidae). Invertebrate Biology.
  • Tyler, P.A., S. Pendlebury, S.W. Mills, L. Mullineaux, K.J. Eckelbarger, M. Baker, C.M. Young. 2008. Reproduction of gastropods from vents on the East Pacific Rise and the Mid-Atlantic Ridge. Journal of Shellfish Research,27:107-118.
  • Eckelbarger, K.J. Oogenesis. 2006. In: Reproductive Biology and Phylogeny of Annelida. B.G.M. Jamieson (ed.), Science Publishers
  • Eckelbarger, K.J. and P. Blades-Eckelbarger. 2005. Oogenesis in calanoid copepods. Invertebrate Reproduction & Development. 47(3):167-182.
  • Eckelbarger, K.J., L. Watling, H. Fournier. 2005. Reproductive biology of the deep-sea polychaete Gorgoniapolynoe caeciliae (Polynoidae), a commensal species associated with octocorals. Journal of the Marine Biological Association of the United Kingdom. 85:1425-1433.
  • Eckelbarger, K.J. 2005. Oogenesis and Oocytes. Hydrobiologia 535/536:179-198.
  • Eckelbarger, K.J. and C.M. Young, 2002. Spermiogenesis and modified sperm morphology in the "Seepworm" Methanoaricia dendrobranchiata (Plychaeta: Orbiniidae) from a methane seep environment in the Gulf of Mexico: implications for fertilization biology. The Biological Bulletin 203:134-143.

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Lewis Incze

Dr. Lewis Incze
Research Professor, School of Marine Sciences
Ph.D. University of Washington, 1983

lincze@maine.com
more info
Download CV

Research interests: Marine ecology; coupling of physical and biological processes in marine population distributions and dynamics; biodiversity, conservation, and ecosystem functioning; ocean observing; climate change; ocean and marine resource management; science and society.

I am interested in the way that ocean physics affects organism distributions, interactions and vital rates such as growth, feeding, and recruitment. Examples of physical processes range from large ocean circulation patterns to eddies and small-scale turbulence, and the organisms I have studied include bivalve mollusks, copepods, larval fish and crustaceans, and whales. I am also interested in systems-level ecology, such as the way that an offshore bank or basin functions (ecologically) by itself and as part of a larger ecosystem. A third area of interest is how this type of knowledge can be used to improve management of human activities in the marine environment, such as fishing, development, and conservation.

  • Jordaan, A, MG Frisk, LS Incze, NH Wolff, L Hamlin, Y Chen. 2013. Multivariate dissemination of species relationships for use in marine spatial planning. Can. J. Fish. Aquat. Sci. 70(2): 316-329, doi: 10.1139/cjfas-2011-0516
  • Lawton, P, LS Incze, SL Ellis. 2012. Representation of biodiversity knowledge within ecosystem-based management approaches for the Gulf of Maine. Amer. Fish. Soc. Sympos. 79: 375-391.
  • Pitcher CR, P Lawton, N Ellis, SJ Smith, LS Incze, C-L Wei, ME Greenlaw, NH Wolff, JA Sameoto, PVR Snelgrove. 2012. Exploring the role of environmental variables in shaping patterns of seabed biodiversity composition in regional-scale ecosystems. Journal of Applied Ecology 2012, 49, 670–679, doi: 10.1111/j.1365-2664.2012.02148.x
  • Ellis, SL, LS Incze, P Lawton, H Ojaveer, BR MacKenzie, CR Pitcher, TC Shirley, M Eero, JW Tunnell Jr., PJ Doherty, BM Zeller. 2011. Four regional marine biodiversity studies: Approaches and contributions to Ecosystem-Based Management. PLoS ONE 6(4)18997 doi:10.1371/ journal.pone.0018997.
  • Johnson, CL, JA Runge, KA Curtis, EG Durbin, JA Hare, LS Incze, J Link, GD Melvin, TD O’Brien, LV Guelpen. 2011. Biodiversity and ecosystem function in the Gulf of Maine: Pattern and role of zooplankton and pelagic nekton. PLoS ONE doi: 10.1371/journal.pone.0016491
  • Li, WKW, RA Andersen, DJ Gifford, LS Incze, JL Martin, CH Pilskaln, JN Rooney-Varga, ME Sieracki, WH Wilson, NH Wolff. 2011. Planktonic microbes in the Gulf of Maine area. PLoS ONE 6(6): e20981. doi:10.1371/journal.pone.0020981
  • Incze, LS, P Lawton, SL Ellis and NH Wolff. 2011. Biodiversity knowledge and its application in the Gulf of Maine Area. Chapter 3 In: (McIntyre, Ed.) Marine Life: Diversity, Abundance and Distribution, Wiley-Blackwell.
  • Fautin, D., P. Dalton, L. S. Incze, J-A. C. Leong, C. Pautzke, A. Rosenberg, P. Sandifer, G. Sedberry, J.W. Tunnell, Jr., I. Abbott, R. E. Brainard, M.Brodeur, L. G. Eldredge, M. Feldman, P. S. Vroom, M. Wainstein, N.Wolff. 2011. An overview of marine biodiversity in United States waters. PloS- One 5(8): e11914 doi: 10.1371/journal.pone.0011914
  • Incze, LS, H Xue, N Wolff, D Xu, C Wilson, R Steneck, R Wahle, P Lawton, N Pettigrew, Y Chen. 2010. Connectivity of lobster populations in the Gulf of Maine. Part II. Coupled biophysical dynamics. Fish. Oceanogr. 19: 1-20, doi  10.1111/j.1365-2419.2009.00522.x
  • Wahle, RA, JS Cobb, LS Incze, P Lawton, M Gibson, R Glenn, C Wilson, J Tremblay. 2010. The American loster settlement index at 20 years: looking back—looking forward. J. Mar. Biol. Assoc. India 52: 180-188.
  • Palumbi, SR, PA Sandifer, JD Allan, MW Beck, DG Fautin, MJ Fogarty, BS Halpern, LS Incze, J-A Leong, E Norse, JJ Stachowitcz, DH Wall. 2009. Managing for ocean biodiversity to sustain marine ecosystem services. Frontiers Ecol. Environment 7, doi: 10.1890/070135
  • Manning, JP, DJ McGillicuddy Jr., NR Pettigrew, JH Churchill, LS Incze. 2009. Drifter observations of the Gulf of Maine Coastal Current. Cont. Shelf Res. 29: 835-845.
  • Xue, H, LS Incze, D Xu, N Wolff, N Pettigrew. 2008. Connectivity of lobster populations in the coastal Gulf of Maine. Part I: Circulation and larval transport potential. Ecol. Model. 210: 193-211.
  • Fogarty, MJ, LS Incze, K Hayhoe, D Mountain, J Manning. 2008. Potential climate change impacts on Atlantic cod (Gadus morhua) off the northeastern USA. Mitig. Adapt. Strat. Global Change. 13:453-466.
  • Stevick, PT, LS Incze, SD Kraus, S Rosen, N Wolff, A Baukus. 2008. Trophic relationships and oceanography on and around a small offshore bank. Mar. Ecol. Progr. Ser. 363: 15-28.

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Pete Jumars

Dr. Pete Jumars
Professor, School of Marine Sciences
Ph.D. Scripps Institution of Oceanography, 1974

jumars@maine.edu
more info
my webpage

Research interests: Benthos, benthic oceanography, biological-physical interactions, bioacoustics

Researchers in my laboratory attempt to identify and quantify important mechanisms by which marine organisms interact with their physical and chemical environments, primarily at the level of individual organisms and primarily but not exclusively in soft-bottom benthic environments. One current project involves the way that diatom chains interact with vortices at the scale of turbulent dissipation. Recent research also involves mechanical interactions of burrowing organisms with mud and sand and the use of acoustics to measure activities, their geological consequences and the abundances and behaviors of animals in and near the seabed. Of long-standing interest are the determinants of the rates at which deposit feeders ingest natural sediments and the nutritional role of various dietary compounds.

  • Jumars, P.A., E. Boss, J.H. Trowbridge and L. Karp-Boss. 2009. Turbulence-plankton interactions: A new cartoon. Marine Ecology: An Evolutionary Approach 30: 233-150.
  • Musielak, M.M., L. Karp-Boss, P.A. Jumars and L.J. Fauci. 2009. Nutrient transport and acquisition by diatom chains in a moving fluid. J. Fluid Mech. 638: 401-421.
  • Clay, T.W., J.B. Fox, D. Grünbaum and P.A. Jumars. 2008. How plankton swim: An interdisciplinary approach for using mathematics and physics to understand the biology of the natural world. Amer. Biol. Teacher 70: 363-370.
  • Dorgan, K.M., S.R. Arwade and P.A. Jumars. 2008. Worms as wedges: Effects of sediment mechanics on burrowing behavior. J. Mar. Res. 66: 219-254.
  • Sato, M., and P.A. Jumars. 2008. Periods and phases of emergence rhythms dependent on season and depth. Limnol. Oceanogr. 53: 1665-1677.
  • Voparil, I.M., L.M. Mayer and P.A. Jumars. 2008. Emulsions vs. micelles in the digestion of lipids by benthic invertebrates. Limnol. Oceanogr. 53: 387-394.
  • Weissberger, E.J., P.A. Jumars, L.M. Mayer and L.L. Schick. 2008. Structure of a Northwest Atlantic shelf macrofaunal assemblage with respect to seasonal variation in sediment nutritional quality. J. Sea Res. 60: 164-175.
  • Dorgan, K.M., S.R. Arwade and P.A. Jumars. 2007. Burrowing in marine muds by crack propagation: kinematics and forces. J. Exp. Biol. 210: 4198-4212.
  • Jumars, P.A. 2007. Habitat coupling by mid-latitude, subtidal, marine mysids: Import-subsidized omnivores. Oceanogr. Mar. Biol. — Ann. Rev. 45: 89-138.
  • Jumars, P.A., K.M. Dorgan, L.M. Mayer, B.P. Boudreau and B.D. Johnson. 2007. Physical constraints on infaunal lifestyles: May the persistent and strong forces be with you. pp. 442-457. In: W. Miller, III, Ed. Trace Fossils: Concepts, Problems, Prospects. Elsevier, Amsterdam.
  • Boss, E., L. Karp-Boss and P.A. Jumars. 2006. Settling of particles in aquatic environments: Low Reynolds numbers. Oceanography 19: 145-148.
  • Dorgan, K.M., P.A. Jumars, B.P. Boudreau and B.D. Johnson. 2006. Macrofaunal burrowing: The medium is the message. Oceanogr. Mar. Biol. – Ann. Rev. 44: 85-121.
  • Abello, H.U., S. M. Shellito, L.H. Taylor and P.A. Jumars. 2005. Light-cued emergence and re-entry events in a strongly tidal estuary, Estuaries 28: 487-499.
  • Boudreau, B.P., I. Croudace, C. Algar, A. Reed, B.D. Johnson, K.M. Dorgan, P.A. Jumars, B.S. Gardiner and Y. Furukawa. 2005. Bubble growth and rise in sediments. Geol. 33: 517-520.
  • Dorgan, K.M., P.A. Jumars, B. Johnson, B.P. Boudreau, and E. Landis. 2005. Burrowing by crack propagation through muddy sediment. Nature 433: 475.
  • Taylor, L.H., S. M. Shellito, H. U. Abello and P.A. Jumars. 2005. Tidally cued emergence events in a strongly tidal estuary, Estuaries 28: 500-509.

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Larry Mayer

Dr. Larry Mayer
Agatha B. Darling Professor of Oceanography, School of Marine Sciences
Ph.D. Dartmouth, 1976

lmayer@maine.edu
more info

Research interests: marine biogeochemistry

We study biogeochemistry - that is, the manner in which organisms and materials (studied chemically) interact in earth surface environments. Major themes are bioavailability of nutritional and toxic materials and implications for carbon and nutrient cycling. We have studied bioavailability to bacteria, phytoplankton and benthic animals, especially as they are affected by interactions with minerals and with sunlight. We consider implications for ecological and biogeochemical cycles - looking at microscopic and organism scales to understand mechanism and at oceanic scales to get at implications for fluxes of materials. Our focus on the ocean is complemented by work in rivers and soils that provide materials brought to the ocean.

  • Chen, Z.* and L. Mayer. 1999. Sedimentary metal bioavailability determined by the digestive constraints of marine deposit feeders: gut retention time and dissolved amino acids. Marine Ecology - Progress Series 176:139-151.
  • Estapa, M.L.* and L.M. Mayer. 2010. Photooxidation of particulate organic matter, carbon/oxygen stoichiometry, and related photoreactions. Marine Chemistry 122:138-147.
  • Kennedy, M., M. Droser, L. M. Mayer, D. Pevear and D. Mrofka. 2006. Late Precambrian Oxygenation; Inception of the Clay Mineral Factory. Science 311:1146-1449.
  • Mayer L. M., R.G. Keil, S.A. Macko, S.B. Joye, K. C. Ruttenberg and R.C. Aller. 1998. The importance of suspended particulates in riverine delivery of bioavailable nitrogen to coastal zones. Global Biogeochemical Cycles 12:573-579.
  • Mayer, L.M. L.L., Schick, K.R. Hardy, and M.L. Estapa*. 2009. Photodissolution and other photochemical changes upon irradiation of algal detritus. Limnology and Oceanography 54:1688-1698.
  • Mayer L., L. Schick, K. Hardy, R. Wagai* and J. McCarthy. 2004. Organic matter content of small mesopores in sediments and soils. Geochimica et Cosmochimica Acta. 68:3863-3872.
  • Mayer, L.M., L.L. Schick, T. Sawyer*, C. Plante, P.A. Jumars, and R.L. Self. 1995. Bioavailable amino acids in sediments: A biomimetic, kinetics-based approach. Limnology and Oceanography 40:511-520.
  • Mayer, L.M., L. Schick, K. Skorko* and E. Boss. 2006. Photodissolution of particulate organic matter from sediments. Limnology and Oceanography 51:1064-1071.
  • Mayer, L.M., D.P. Weston and M.J. Bock. 2001. Benzo-a-pyrene and zinc solubilization by digestive fluids of benthic invertebrates - a cross-phyletic study. Environmental Chemistry and Toxicology 20:1890-1900.
  • Voparil, I.M.* and L.M. Mayer. 2004. Commercially-available chemicals that mimic a deposit-feeder's (Arenicola marina) digestive solubilization of lipids. Environmental Science and Technology 38:4334 - 4339.

* - students
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Mary Jane Perry

Dr. Mary Jane Perry
Professor, School of Marine Sciences
Director, Darling Marine Center
Ph.D. Scripps Institution of Oceanography, 1974

perrymj@maine.edu
more info
Phytoplankton & Optics Lab
Damariscotta River Chlorophyll data - daily dock sampling

Research interests: biological oceanography and biological optics

My long-term goal is to understand the mechanisms responsible for the variability in phytoplankton biomass, primary production, and species composition. Though I started my oceanographic career by studying the role of nutrients in controlling phytoplankton biomass and production in the subtropical Central North Pacific, my current focus is on the interaction of phytoplankton and light in the ocean. I started this phase of my career with a study of the photoadaptive changes in the absorption cross section of photosystem I in marine phytoplankton. My present research interests include primary production (at the level of the single cell and as well as the entire phytoplankton assemblage); photosynthetic physiology as well as phytoplankton physiology in general; biological optics; and ocean observations using in-situ optics and remote sensing. I have participated in a number of cruises in both the north Atlantic and Pacific oceans.

  • Boss, E., M.J. Perry, and M.C. Talbot. 1999. Observation of an intense deep-water intrusion in Puget Sound. Puget Sound Research ’98 Conference Proceedings.
  • Culver, M.E., and M.J. Perry. 1997. Calculation of solar-induced fluorescence from surface and subsurface waters. Journal of Geophysical Research 102:10,563-10,572.
  • Culver, M.E., and M.J. Perry. 1999. Fluorescence excitation estimates of photosynthetic absorption coefficients for phytoplankton and their response to irradiance. Limnology and Oceanography 44: 24-36.
  • English, D.C., K. Banse, D.L. Martin, and M.J. Perry. 1996. Electronic overshoot and other bias in the CZCS global data set: comparison with ground truth from the Subarctic Pacific. Journal of Remote Sensing 17:3157-3168.
  • Lee, Z.P., K.L. Carder, J. Marra, R.G. Steward, and M.J. Perry. 1996. Estimating primary production at depth from remote sensing. Applied Optics 35:463- 474.

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Warren Riess

Dr. Warren Riess
Research Associate Professor, School of Marine Sciences
Research Associate Professor, Department of History
Ph.D. University of New Hampshire, 1987

riess@maine.edu
more info
Penobscot Expedition 1779

Research interests: maritime history and archeology

My general area of interest is the Atlantic community during the fifteenth through eighteenth centuries, especially North America and the Caribbean. My research interests focus on the history and archaeology of the early American merchant trade and navy. Within these spheres I use a combined approach of economic, military, political, social, and technical history. Major areas of current research are the archaeological sites of two eighteenth-century merchant ships: the Nottingham Galley off Boon Island, Maine and the Ronson ship in Manhattan. My archaeological expertise is in the design and construction of early vessels and electronic search and survey.

  • Riess, W.C., “Nottingham Galley at Boon Island: the History, Archaeology, and Novel,” The Northern Mariner/le marin du nord, XVIII, 2008, 21-26.
  • Riess, W.C., “Maine, the First Twenty-five Years,” Handbook of American Maritime Archaeology, ed. J. Barstad, Plenum Press, 2002, 29-41.
  • Riess, W.C. The Angel Gabriel: The Elusive English Galleon. Maine: 1797 House, 2001.
  • Riess, W.C., "Penobscot Expedition" and "Ronson Ship," British Museum Encyclopedia of Maritime and Underwater Archaeology, ed. J. Delgado, Yale University Press, 1997.
  • Riess, Warren and Geoffrey Daniel, "Evaluation of Preservation Efforts for the Revolutionary War Privateer Defence," International Journal of Nautical Archaeology, 26.4, 1997, 330-338.

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Bob Steneck

Dr. Bob Steneck
Professor, School of Marine Sciences
Pew Fellow in Marine Conservation
Ph.D. Johns Hopkins University, 1982

steneck@maine.edu
Full CV
more info

Research interests: ecology, evolutionary biology, fisheries oceanography, plant-herbivore interactions

I study ecological processes in the benthic marine realm. My experiments focus on the food webs, structure and dominant organisms of coastal marine communities. My "laboratory" is the subtidal zone in which I use SCUBA diving, underwater video systems, manned submersibles and operated vehicles as research tools to study the Gulf of Maine and Caribbean. My current major research projects focus on the ecology of the settling corals, the impact of large predatory and herbivorous fishes on benthic marine communities and plant-herbivore interactions from both the botanical and zoological perspectives. In the Gulf of Maine, my work focuses on applying ecosystem based management to fisheries and in studying the potential environmental impacts of offshore wind energy. In the Caribbean I'm studying how herbivory affects the recruitment of corals and the resilience of coral reef ecosystems.

  • Estes, J.A., C.H. Peterson, and R.S. Steneck. 2010. Some effects of apex predators in higher-latitude coastal oceans.  Pages  37 – 53 In: Trophic Cascades: predators, prey, and the changing dynamics of nature.  Terborgh, J. and Estes, J. S. (eds).  Island Press, Washington, DC.
  • Incze, L., H. Xue, N. Wolff, D. Xu, C. Wilson and R.S. Steneck. 2010. Connectivity of lobster populations in the coastal Gulf of Maine Part II. Coupled biophysical dynamics. Fisheries Oceanography. Fish. Oceanogr. 19:1, 1–20.
  • Steneck R.S., and J.A. Wilson. 2010. A fisheries play in an ecosystem theater: challenges of managing ecological and social drivers of marine fisheries at nested spatial scales. Bulletin of Marine Science 86(2): 387–411.
  • Arnold, S., P. Mumby and R.S.Steneck. 2010. Running the gauntlet to coral recruitment through a sequence of local multiscale processes. Marine Ecology Progress Series. DOI 10.3354/meps08724.
  • Foley, M., B.S. Halpern, F. Micheli, M.H. Armsby, M.R. Caldwell, E. Prahler, D. Silvas, C. Crain, M. Rohr, M.W. Beck, M.H. Carr, L.B. Crowder, E. Duffy, S. Hacker, K. McLeod, C.H. Peterson, H.M. Regan, P.A. Sandifer, R.S. Steneck.  2010. Guiding scientific principles for marine spatial planning. Marine Policy. doi:10.1016/j.marpol.2010.02.001
  • 2010 Gelcich, S., Hughes, T. P., Olsson, P., Folke, C., Defeo, O., Fernandez, M., Foal, S., Gunderson, L. H., Rodriguez-Sickert, C., Scheffer, M., Steneck, R., and Castilla, J. C. Navigating transformations in governance of Chilean marine coastal resources.  Proc. Of Nat. Acad. Sciences.  Doi: 10.1073/pnas.1012021107
  • Lorenzen, K., R. Steneck, R.R. Warner, A.M. Parma, F.C. Colemen and K.M. Leber. 2010. The spatial dimensions of fisheries: putting it all in place.  An introduction to the proceedings of the seventh William R. and Lenore Mote Symposium in Fisheries Ecology.  Bulletin of Marine Science. 86(2): 169–177
  • Wells, R.J.D., R.S. Steneck, A.T. Palma. 2010., Three-dimensional resource partitioning between American lobster (Homarus americanus) and rock crab (Cancer irroratus) in a subtidal kelp forest, J. Exp. Mar. Biol. Ecol. (2010), doi:10.1016/j.jembe.2010.01.008
  • Hughes, T., N. Graham, J. Jackson, P. Mumby, R. Steneck. 2010. Rising to the challenge of sustaining coral reef resilience.  Trends in Ecology and Evolution.  doi: 10.1016/j.tree.2010.07.011
  • Williams, B., J.Halfar, R.S. Steneck, U.G. Wortmann, S. Hetzinger, W. Adey, P. Lebednik, and M. Joachimski. 2010. Twentieth century δ13C variability in surface water dissolved inorganic carbon recorded by coralline algae in the northern North Pacific Ocean and the Bering Sea, Biogeosciences Discuss., 7, 5801-5828, doi:10.5194/bgd-7-5801-2010
  • Mumby, P.J. and R.S. Steneck. 2011. The resilience of coral reefs and its implications for reef management.  Chapter 29 in. Dubinsky, Z. and Stambler, N. (eds) Coral Reefs: An Ecosystem in Transition. Springer Press. DOI 10.1007/978-94-007-0114-4_29.

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Rick

Dr. Rick Wahle
Research Professor, School of Marine Sciences
Ph.D., Zoology, University of Maine, 1990

richard.wahle@maine.edu
more info
Wahle Lab Webpage

Research Interests: benthic ecology, fisheries science

I am broadly interested in the influences of the ocean's physical and biotic environment on the population dynamics and distribution of marine benthic organisms. Much of my research integrates the fields of ecology, oceanography, and fishery science toward a better mechanistic understanding of marine populations and communities. I have a particularly long-standing research program on the American lobster for which I and my collaborators are striving to develop predictive tools for population trends through an understanding of larval transport, settlement, and post-settlement processes.

  • Geraldi, N.R.*, Wahle, R.A. Dunnington, M.J. 2009. Habitat effects on American lobster catch and movement: Insights from geo-referenced trap arrays, seabed mapping, and tagging. Can. J. Fish. Aquat. Sci. 66: 460-470.
  • Wahle, R.A., M. Gibson, M.J. Fogarty. 2009. Distinguishing disease impacts from larval supply effects in a lobster fishery collapse. Mar. Ecol. Prog. Ser. 376: 185–192. doi: 10.3354/meps07803 (Selected as Editors’ Choice in Science 323:1406b; 3/13/09)
  • Wahle, R.A., C. Bergeron*, C. Wilson, M. Parkhurst. 2009. A vessel-deployed passive post-larval collector for the American lobster. New Zealand J. Mar. Freshw. Res. 43: 465 -474.
  • Wahle, R.A., C.E. Bergeron*, A. Chute, Y. Chen, L. Jacobson. 2008. Northwest Atlantic deep-sea red crab before and after the onset of harvesting. ICES J. Mar. Sci. 65: doi:10.1093/icesjms/fsn058
  • Frumhoff, P.C., J.J. McCarthy, J.M. Melillo, S.C. Moser, and D.J. Wuebbles. 2007. Confronting Climate Change in the U.S. Northeast: Science, Impacts, and Solutions. Synthesis report of the Northeast Climate Impacts Assessment (NECIA). Cambridge, MA: Union of Concerned Scientists (UCS). (Member of contributing team of white paper authors)
  • Selgrath, J., K*. Hovel, R. Wahle. 2007. Effects of habitat edges on American lobster abundance & survival. J. Exp. Mar. Biol. Ecol. 353: 253–264.
  • O’Donnell, K.P.*, Wahle, R.A., Dunnington, M.J., M. Bell. 2007. Spatially referenced trap arrays detect sediment disposal impacts in a New England estuary. Mar. Ecol. Prog. Ser. 348: 249–260.

* Student authors
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Rhian

Dr. Rhian Waller
Associate Research Professor
Ph.D. Southampton Oceanography Center, UK, 2004

rhian.waller@maine.edu
more info
Waller Lab Webpage

Research Interests: Invertebrate reproduction and larval development, deep-sea biology, cold-water corals, biogeography, larval dispersal and population connectivity

My research interests center on the reproduction and development of cold-water and deep-sea invertebrates from around the globe, and how these animals are affected by both natural and anthropogenic environmental change. My laboratory has projects from the Arctic to the Antarctic and many places in between. Though we primarily use microscopy techniques (histology, SEM and TEM) we also use geographic databases and genetic techniques to examine biogeographic patterns of species dispersal and make hypotheses on larval transport mechanisms. I welcome requests from undergraduates seeking laboratory experience, as well as students thinking about graduate school. Photo credit - Dann Blackwood, USGS.

  • Bayer, S., L.S. Mullineaux, R.G.Waller& A.R. Solow. 2010.Reproductive traits of pioneer gastropod species colonizing deep-sea hydrothermal vents after an eruption. Marine Biology, DOI: 10.1007/s00227-010-1550-1.
  • Scanlon, K.M., R.G. Waller, A. Sirotek, J.M. Knisel & S.M. Alesandrini. 2010. USGS Cold-Water Coral Geographic Database – Gulf of Mexico and Western North Atlantic, Version 1.0. USGS Open File Report
  • Waller, R.G. & P.A. Tyler. 2010. Reproductive patterns in two deep-water solitary corals from the North-east Atlantic—Flabellum alabastrum and F. angulare (Cnidaria: Anthozoa: Scleractinia). Journal of the Marine Biological Association, UK.
  • Waller, R.G., P.A. Tyler & C. Smith. 2008. Fecundity and embryo development of three Antarctic deep-water scleractinians: Flabellum thouarsii, F. curvatum and F. impensum. Deep-Sea Research II, 55: 2527-2534.
  • Flint, H., R.G. Waller & P.A. Tyler. 2007. Reproduction in Fungiacyathus marenzelleri from the northeast Pacific Ocean. Marine Biology, 151(3): 843-849.
  • Waller, R.G., J. Adkins, L. Robinson & T.M. Shank. 2007. Ancient DNA techniques and their utility in deep-water coral research. Bulletin of Marine Science, 81(3): 351-359.
  • Robinson, L.F., J.F. Adkins, D.S. Scheirer, D.F. Fernandez, A. Gagnon & R.G. Waller. 2007. Deep-sea scleractinian coral age and depth distributions in the NW Atlantic for the last 225 thousand years. Bulletin of Marine Science, 81(3): 371-391.
  • Waller, R.G. & A. Baco-Taylor. 2007. Reproductive morphology of three Hawaiian deep-water precious corals. Bulletin of Marine Science, 81(3): 533-542.
  • Waller, R.G., L. Watling, P. Auster & T.M. Shank. 2007. Fishing Impacts on the Corner Rise Seamounts. Journal of the Marine Biological Association, UK, 87: 1075-1076.
  • Waller, R.G. 2005. Deep Water Scleractinians: Current knowledge of reproductive processes. In: Freiwald A & Roberts JM (eds) Cold-water Corals and Ecosystems. Springer, Heidelberg, 691-700.

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Les Watling

Dr. Les Watling
Professor Emeritus, School of Marine Sciences
Pew Fellow in Marine Conservation
Ph.D. Delaware, 1974

watling@maine.edu
more info
Watling Lab Webpage

Research interests: benthic ecology, taxonomy

Dr. Watling's research interests span two disparate topics: crustacean taxonomy and phylogeny and benthic oceanography. His crustacean interests center on amphipods and cumaceans, but he has studied a variety of other groups, including spelaeogriphaceans and syncarids. These two groups are southern hemisphere phylogenetic relicts about which little is known. Dr Watling's benthic interests are focused on impacts of humans on benthic environments, with an emphasis on organic enrichment and habitat disruption. Topics investigated in the last few years include the impact of salmon net-pen aquaculture on benthic environments, and the effects of fishing activities on benthic habitats and its consequences for benthic community structure.

  • Watling, L., Haedrich, R.L., Devine, J., Drazen, J., Dunn, M.R., Gianni, M., Baker, K., Cailliet, G., Figueiredo, I., Kyne, P.M., Menezes, G., Neat, F., Orlov, A., Duran, P., Perez, J.A., Ardron, J.A., Bezaury, J., Revenga, C., and Nouvian, C. 2011. Can ecosystem-based deep-sea fishing be sustained? Report of a workshop held 31 August-3 September 2010. Walpole, ME: University of Maine, Darling Marine Center. Darling Marine Center Special Publication 11-1. 84p.
  • Gerken, S. & L. Watling. 1999. Cumacea (Crustacea) of the Faroe Islands Region. Fró_skaparrit 47: 199-227.
  • Gerken, S., L. Watling, & A.B. Klitgaard. 2000. Contumacious Beasts: A story of two Diastylidae (Cumacea) from Arctic waters. Journal of Crustacean Biology 20 (1) 31-43.
  • Gerken, S., L. Watling, A. Klitgaard. 2000. Some contumacious Cumacea from Arctic waters. Journal of Crustacean Biology 20: 31-43.
  • Hessler, R.R. & L. Watling. 1999. Les Péracarides: un groupe controversé. Traité de Zoologie, Anatomie, Systematique, Biologie. Tome VII, Fascicule III A, Crustacés Péracarides. Mémoires de l'Institut Océanographique, Monaco, 19: 1-10.
  • Johnson, W.E., P. Stephens & L. Watling. 2000. Reproductive biology of peracarid crustaceans. Advances in Marine Biology, 39:105-260.
  • Norse, E. & L. Watling. 1999. Impacts of mobile fishing gear: the biodiversity perspective. Pp. 31-40, in L. Benaka, editor. Fish habitat: essential fish habitat and rehabilitation. American Fisheries Society Symposium 22, Bethesda, Maryland.
  • Petrescu, I. & L. Watling. 1999. Revision of genus Americuma Watling, 1991 (Crustacea, Cumacea) with the description of Styloptocuma heardi (Bacescu, 1979) based on neotype material. Travaux du Museum National d'Histoire naturelle "Grigore Antipa", 41: 299-308.
  • Watling, L. & S. Gerken. 1999. Two new cumacean (Crustacea) species from the deep South Atlantic. Zoosystema, Paris, 21(4): 661-669.
  • Watling, L. 1999. Toward understanding the relationships of the peracaridan orders: the necessity of determining exact homologies. Pp. 73-89, In F.R. Schram & J.C. von Vaupel Klein (editors). Crustaceans and the Biodiversity Crisis. Proceeding of the Fourth International Crustacean Congress, Amsterdam. Brill, Leiden, The Nether lands.
  • Watling, L. and S. Gerken. 1999. Leucon (Crymoleucon) noerrevangi, a new species of leuconid (Cumacea: Crustacea) from the Faroe Islands. Sarsia 84(5) 437-444.
  • Watling, L., F.R. Schram & C.H.J. Hof. 2000. The place of the Hoplocarida in the malacostracan pantheon. Journal of Crustacean Biology 20 (Special Number 2): 1-11.

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